Actins are highly conserved proteins that are involved in various types of cell motility and are ubiquitously expressed in all eukaryotic cells. As an essential component of cell cytoskeleton, actin plays an important role in cytoplasmic streaming, cell shape determination, cell division, organelle movement and extension growth. In plants, actin preferentially expressed in young and expanding tissues, floral organ primordia, developing seeds, and emerging inflorescence. It displays a very high expression level in mature pollen and pollen tubes.